Commentary for Keritot 4:50
שאם עשאן כולן בהעלם אחת חייב על כל אחת ואחת
all other forbidden relations are thus made analogous to 'his sister': Just as in the case of 'his sister' one is liable on her account to a separate offering, so als other cases one is liable to a separate offering for each [transgression].
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